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Dryinus Latreille

(Life: Kingdom: Metazoa (animals); Phylum: Arthropoda; Class: Hexapoda; Order: Hymenoptera; Superfamily: Chrysidoidea; Family: Dryinidae; Subfamily: Dryininae)

Dryinus Latreille, 1804. Type species: Dryinus formicarius Latreille, 1804, by subsequent monotypy (Latreille 1805).

Bocchoides Benoit, 1953. Type species: Bocchoides bekilyensis Benoit, 1953.

ClassificationIdentification keys


Drynis aberrans Benoit, 1954

Dryinus aethiopicus (Olmi, 1984)


Dryinus afer (Olmi, 1989)


Dryinus alticolus (Benoit, 1951)

Dryinus basilewskyi Benoit, 1953

  Dryinus bekilyensis (Benoit, 1954)

Dryinus bellicosus Olmi, Copeland & van Noort, 2019

  Dryinus berlandi Benoit, 1954 (incertae sedis)
  Dryinus botswanensis (Olmi, 1991)

Dryinus burgeoni Benoit, 1953


Dryinus chamaeleo Benoit, 1953

  Dryinus cariniceps Cameron, 1906
Dryinus copelandi Olmi, 2011
  Dryinus daviesi Olmi, 2009
  Dryinus dayianus Olmi, 1993

Dryinus deceptor (Turner, 1928)

Dryinus dentatiforceps Olmi, Copeland & van Noort, 2019

Dryinus digo Olmi, Copeland & Guglielmino, 2015

Dryinus eardleyi (Olmi, 1987)

Dryinus erenianus Olmi, Copeland & van Noort, 2019


Dryinus erraticus Turner, 1928

  Dryinus evertsi Olmi, 1989

Dryinus grobbelaari Olmi 2007

  Dryinus grossus Olmi, 2002
  Dryinus hararianus Olmi, 1987

Dryinus hova (Ceballos, 1936)

  Dryinus irwini Olmi, 2007
  Dryinus ivoriensis (Olmi, 1989)
Dryinus kibalus Olmi, van Noort & Guglielmino, 2016
  Dryinus luweli (Benoit, 1951)
  Dryinus madagascolus Olmi, 1993

Dryinus mahensis (Kieffer, 1912)


Dryinus mayogo (Benoit, 1953)

Dryinus milleri Olmi, Copeland & van Noort, 2019

Dryinus mobotensis Olmi, Copeland & van Noort, 2019


Dryinus modestus Olmi, 1984

Dryinus nigrithorax Olmi, Copeland & van Noort, 2019

  Dryinus noyesi (Olmi, 1987)

Dryinus orophilus (Benoit, 1950)

  Dryinus paulyi Olmi, 1989
  Dryinus praecipuus Olmi, 2002

Dryinus pretorianus Olmi, 1984

  Dryinus pseudoafer Olmi, 1993

Dryinus saussurei (Ceballos, 1936)


Dryinus schoutedeni (Benoit, 1953)


Dryinus seyrigi Benoit, 1953

  Dryinus shimbanus Olmi, 2011

Dryinus sierranus Olmi, 1984

Dryinus spangleri Olmi, 1984

Dryinus teres Olmi, Copeland & van Noort, 2019

Dryinus tulearensis Olmi, Copeland & van Noort, 2019


Dryinus turneri Olmi, Copeland & Guglielmino, 2015


Dryinus ugandanus (Olmi, 1984)


Dryinus undulatus Benoit, 1950

  Dryinus vannoorti Olmi, 2006
Dryinus whittleorum Olmi, Copeland & van Noort, 2019
  Dryinus yemenensis Olmi & van Harten, 2006
  Dryinus zambeziacus Olmi, 2004

Dryinus species




Hosts: Dictyopharidae, Cixiidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Fulgoridae.


Benoit, P.G.L.1951. Exploration du Parc National Albert, Mission G. F. de Witte (1933-1935). Fasc. 73. Dryinidae (Hymenoptera Aculeata), Evaniidae (Hymenoptera Terebrantia). Hayez, Bruxelles, 26pp.

Benoit, P.G.L. 1953. Nouveaux Dryinidae (Hym.) du continent africain. Ann. Mus. Civ. St. Nat. Genova 64: 298-302.

Benoit, P.L.G., 1954. Monographie des Dryninides Malgaches (Hym.-Acul.). Mém Inst. Sci. Madagascar, Sér. E, IV:383-430.

Cameron P., 1906. Description of a new species of Dryinus from Cape Colony (Hym.). Zeitschr. Hymenopterol 3:158-159.

Ceballos, G., 1936. Estudios sobre Anteoninae de Madagascar (Hym. Bethyl.). EOS 12: 43-64.

Guglielmino, A. & Olmi, M. 1997. A host-parasite catalog of world Dryinidae (Hymenoptera: Chrysidoidea). Contributions on Entomology, International 2 (2): 165–298.

Guglielmino, A. & Olmi, M. 2006. A host-parasite catalog of world Dryinidae (Hymenoptera: Chrysidoidea): first supplement. Zootaxa 1139: 35–62.

Guglielmino, A. & Olmi, M. 2007. A host-parasite catalog of world Dryinidae (Hymenoptera: Chrysidoidea): second supplement. Bollettino di Zoologia Agraria e di Bachicoltura, Ser. II 39 (2): 121–129.

Guglielmino A, Olmi M, Bückle C. 2013. An updated host-parasite catalogue of world Dryinidae (Hymenoptera: Chrysidoidea). Zootaxa 3740: 1–113. doi: 10.11646/zootaxa.3740.1.1

Kieffer, J.J. 1912. Hymenoptera Proctotrupoidea. In Percy Sladen Trust Expedition, Trans, Linn. Soc. London 15: 45-80.

Kieffer J.J. & Marshall, T.A. 1906. Proctotrypidae. In André E., Species des Hyménoptères d'Europe et d'Algerie 9: 289-552.

Latreille, P.A., 1804. Nouvelle dictionnaire d'Histoire naturelle 24, Paris.

Moczar L., 1965. Remarks of some od Drynini and Gonatopodini (Hymenoptera). Ann. Hist.-Nat. Mus. Nation. Hung. 57: 375-406.

Olmi, M. 1975. Paradryinus afer sp. n., primo Driinde parassita degli Omotteri Fulgoroidei Ricaniidi del genere Ricania Germ. (Hymenoptera, Dryinidae). Boll. Ist. Ent. Univ. Bologna32: 221-226.

Olmi, M. 1984. A Revision of The Dryinidae (Hymenoptera). Memoirs of the American Entomological Institute, 37: 1-947.

Olmi, M. 1987. Descrizone di nuove specie di Dryinidae (Hymenoptera Chrysidoidea). Boll. Zool. agr. Bachic., Ser. II, 19: 31-70.

Olmi, M. 1987. New Species of Dryinidae (Hymenoptera Chrysidoidea). Fragm. Entomol., Roma, 19: 371-456.

Olmi, M. 1989. New Dryinidae from Africa and Asia (Hymenoptera Chrysidoidea). Oriental Insects, Vol, 23: 157-162.

Olmi, M. 1989. Supplement to the Revision of the World Dryinidae (Hymenoptera Chrysidoidea). Frustula Entomologica, Nuova Serie, vol. 12: 109-395.

Olmi, M. 1993. A new generic classification for Thaumatodryininae, Dryininae and Gonatopodinae, with descriptions of new species (Hymenoptera Dryinidae). Boll. Zool. agr. Bachic., Ser. II, 25: 57-89.

Olmi, M. 2002. New Species of Dryinidae and Embolemidae from Madagascar (Hymenoptera Chrysidoidea). Frustula entomol., Nuova Serie 25: 86-109.

Olmi, M. 2004. A Contribution to the Knowledge of Dryinidae (Hymenoptera Chrysidoidea) of Northern Mozambique. Oriental Insects, Vol. 38: 353-372.

Olmi, M. 2006. A catalogue of Dryinidae and Embolemidae of South Africa, with descriptions of new species (Hymenoptera Chrysidodea). Frustula entomologica (2005)(N.S.) vol. 28-29 (41-42): 1-57.

Olmi, M. 2007. New species of Afrotropical Dryinidae (Hymenoptera: Chrysidoidea), with description of a new genus and a new subfamily. African Invertebrates, vol  48: 199-232.

Olmi, M. 2011. A contribution to the knowledge of world Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica (2009) (N.S.) XXXII (XLV): 43-76.

Olmi, M. & Copeland, R.S. 2011. A catalogue of Dryinidae, Embolemidae and Sclerogibbidae of Kenya and Burundi, with descriptions of new species (Hymenoptera: Chrysidoidea). African Invertebrates 52: 177–206.

Olmi, M. &  van Harten, A. 2006. Dryinidae, Sclerogibbidae and Embolemidae (Hymenoptera: Chrysidoidea) of Yemen, with revised keys to the species of the Arabian peninsula. Fauna of Arabia 21: 307–337.

Olmi, M. & Copeland, R.S. & Guglielmino, A. 2015. An updated checklist of Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera) of Kenya and Burundi, with descriptions of thirteen new species. Acta Entomologica Musei Nationalis Pragae 55(1): 333-380.

Olmi, M., Copeland, R.S. & van Noort, S. 2019. Dryinidae of the Afrotropical region (Hymenoptera: Chrysidoidea). Zootaxa 4630: 1–619. http://dx.doi.org/10.11646/zootaxa.4630.1.1

Olmi, M., van Noort, S. & Guglielmino, A. 2016. Contribution to the knowledge of Afrotropical Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera) with description of new species from Central African Republic and Uganda. ZooKeys 578: 45-95.

Turner, R. E., 1928. New Hymenoptera of the Family Bethylidae. Ann. Mag. Nat. Hist., Ser. 10, Vol. 1: 129-152.


Photographs © Charissa de Lange (Pretoria) or © Simon van Noort (Iziko Museums of South Africa) or © Bob Copeland (ICIPE).

Web author Simon van Noort (Iziko South African Museum)


Citation: van Noort, S. 2022. WaspWeb: Hymenoptera of the Afrotropical region. URL: www.waspweb.org (accessed on <day/month/year>).

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